4.1. Assess the conservation status of all species and identify those needing targeted recovery actions.
Knowledge of the distribution, abundance, trends, and threats provides an essential baseline for conservation planning and action. The primary tools for assessing the status of species are the IUCN Red List of Threatened Species, the Green Status of Species, and the Living Planet Index. Species monitoring is supported by several technical tools.
Subactions
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4.1.1. Assess the extinction risk, population size and trends, distribution, threats, and conservation potential of all species. - esIUCN Red List PartnersIUCN SSC Specialist Groups and Red List AuthoritiesWorld Widelife Fund for NatureInternational Whaling CommissionInternational Treaty on Plant Genetic Resources for Food and AgricultureConvention on the Conservation of Migratory Species of Wild AnimalsReverse the Red partnership
Primary tools and resources
The Global Strategy for Plant Conservation (GSPC)
The GSPC – A Plan to Save the World’s Plant Species – highlights the importance of plants and the ecosystem services they provide for all life on earth, and aims to ensure their conservation. The GSPC includes 16 targets for plant conservation. An updated GSPC was formally adopted at the 10th meeting of the Conference of the Parties to the CBD.
Living Planet Index (LPI)
The Living Planet Index measures the state of the world’s biodiversity based on population trends of vertebrate species from terrestrial, freshwater, and marine habitats. The LPI has been adopted by the Convention of Biological Diversity (CBD) as an indicator of progress towards its targets. The LPI is compiled by WWF and ZSL based on trends in thousands of population time-series data collected from monitored sites around the world. The Living Planet Database contains population time-series data on the abundance of tens of thousands of vertebrate species between 1970 and 2021. It is publicly available, except for some confidential records that cannot be shared.
How to use
- The annual Living Planet Report can be downloaded at: https://www.wwf.org.uk/our-reports/living-planet-report
- The latest results and the technical supplement are available at: https://www.livingplanetindex.org/
- Data in the Living Planet Database can be searched or downloaded at:
Species Threat Abatement and Restoration (STAR) metric
The STAR metric assesses the potential of particular actions at a specific location to contribute to reduce species extinction risk / global targets for species. It measures the potential contribution of two kinds of action: threat abatement / reduction and habitat restoration using data on the distribution, threats, and extinction risk of threatened species contained in the IUCN Red List. It helps governments, the finance industry, investors, and companies to target their investments and activities to achieve conservation outcomes for threatened species. The STAR metric can be applied to any location. It is maintained under the authority of the IUCN Red List Committee.
IUCN Red List of Threatened Species
The IUCN Red List of Threatened Species (the Red List) is the globally recognised standard on assessing extinction risk. Red List assessments are used by the Convention on Biological Diversity, CITES, other inter-governmental agreements, national governments, and conservation planners. The Red List assigns species into one of 8 categories based on 5 criteria, all with quantitative thresholds. The three highest categories of threat – Critically Endangered, Endangered, Vulnerable – are collectively considered ‘Threatened’. The Red List process has been formalized to support objectivity and scientific rigour. Red List assessments are carried out by SSC Red List Authorities, Red List Partners, IUCN staff, and regional and national agencies. In addition, Red List assessments provide a compendium of information on species, (taxonomy, geographic range, population, habitat and ecology, threats, conservation actions). The Red List operates at global, regional, and national levels. Guidelines for application at national or regional levels have been developed and the National Red List Working Group provides additional support.
Red List Authorities have been established for all major taxonomic groups included on the IUCN Red List. In most cases, the RLA is a sub-group within an IUCN SSC Specialist Group responsible for the species, groups of species or species within a specific geographic area. There are some exceptions; for example, BirdLife International is the designated RLA for all birds.
The process is guided by Rules of Procedure, supported by guidelines on application of the Red List Categories and Criteria, the Species Information Service (SIS) data entry and management system, and certification for assessors and RL trainers. All these materials can be downloaded, free, from the Red List website. A free, online training course for Red List assessors, is available.
Species can be assessed for the Red List at global, regional, and national levels. Guidelines for the Regional (and national) Application of the Categories and Criteria have been developed and support for development of national Red Lists is available from the National Red List Working Group.
How to use
- To check the global Red List status of a species and access the supporting information, visit the species assessment page on the Red List website
- To download the Red List Categories and Criteria, Guidelines, Rules of Procedure, and Guidelines for Application at Regional Level go the Red List resources
- To initiate or to contribute to a global assessment, contact the Chair of the relevant IUCN SSC Specialist Group or Red List Authority, for birds, see BirdLife International
- For taxonomic groups not listed, contact the relevant IUCN SSC Conservation Committee: Marine, Invertebrate, Plant, Fungi, Freshwater, Red List, Standards and Petitions.
- To develop a National Red List for a species or group of species visit the National Red List Working Group
- To obtain details or register for an online Red List training course: https://www.iucnredlist.org/resources/online
IUCN Green Status of Species
The decline of many species towards extinction has largely focused conservation efforts on ensuring that species remain extant. However, conservationists have long recognised the need to complement this by aiming to recover depleted populations throughout a species’ range and to restore species to ecosystems from which they have been extirpated. The main objectives of the IUCN Green Status of Species are: to provide a standardised framework for measuring species recovery; to recognise conservation achievements; to highlight species whose current conservation status is dependent on continued conservation actions; to forecast the expected conservation impact of planned conservation action; and to elevate levels of ambition for long-term species recovery. These objectives together encourage conservation towards species recovery, throughout a species’ range.
IUCN Red List index : guidance for national and regional use. Version 1.1
The RLI monitors trends in extinction risk over time and is based on the IUCN Red List of Threatened Species. It tracks the conservation status of major species groups, and has been applied so far to birds, mammals, cycads, amphibians, corals. The data can be disaggregated by region, or country.
How to use
For details of the RLI process and to see results: https://www.iucnredlist.org/assessment/red-list-index
Other tools and resources
Species Threat Abatement and Restoration in the Central African Republic
This report presents findings from an assessment of the biodiversity conservation potential of four project sites: Lobaye, Mambéré-Kadéï, Ombella-Mpoko, and Sangha-Mbaéré located in the Bangui region of the Central African Republic (CAR). This assessment was done using the Species Threat Abatement and Recovery (STAR) metric, which employs high-resolution imagery and an approach to modelling species’ Area of Habitat (AOH) that was revised after June 2020 among other enhancements. This work was undertaken to better inform threat abatement and restoration planning and implementation at the partner project sites, and as part of a wider effort to pilot and strengthen the use of STAR as a tool for restoration and conservation practitioners, communities, investors, and policymakers.
Conservation actions in the national ITPGRFA Reports
These are the country reports on implementation of the International Treaty received from Contracting Parties pursuant to Section V.1 of the “Procedures and operational mechanisms to promote compliance and address issues of non-compliance” (Resolution 9/2013), for which an updated Standard Reporting Format was adopted by the Governing Body in 2019 (Resolution 7/2019).
The Compliance Committee prepares a comprehensive synthesis and succinct analysis report for each session of the Governing Body. Please see the latest version here.
Some of the information received from Contracting Parties through their national reports is reused under several indicators in the framework of the Sustainable Development Goals (SDGs) reporting process. See, for example, SDG Indicator 15.6.1.
The deadline for the submission of national reports under the Second Reporting Cycle was extended by the Governing Body until 1 October 2024.
IWC Extinctions Initiative
According to the IUCN Red List a species or subspecies is considered Extinct (EX) when there is no reasonable doubt that the last individual has died. A species or subspecies is presumed Extinct when exhaustive surveys in known and expected habitat have failed to record a single individual.
Distinct populations can also become locally extinct. A single species is often made up of several separate populations and one population may be eradicated whilst others continue to live. For example, the North Pacific gray whale population in the eastern North Pacific is considered healthy, but the western population is critically endangered.
Extinction is a natural phenomenon, but current rates of extinction are much higher than historic ‘background rates.’ The lifespan of a species varies depending on a number of factors such as body size and geographic range. Many cetacean species are assessed to have survived between 5 and 30 million years. Today’s increased rates of extinction are due to human activities and associated habitat disruption and climate change.
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, assesses the conservation status of most plant and animal species, some subspecies, and some subpopulations. They use a rigorous approach to assess all available data for each species against defined criteria, and classify each into one of the IUCN Red List status categories
ThreatSearch
ThreatSearch is a global database of all known conservation assessments of plants. It contains global, regional and national conservation assessments for plants from a variety of sources. The aim is for ThreatSearch to be a one-stop shop to find any conservation assessment for plants.
Together with our two main collaborators – the National Red List and the Royal Botanic Gardens, Kew – we have assembled currently available conservation assessments into a single list of conservation assessments for plants. Additional significant contributors include NatureServe and CNCFlora. We are continuing to add new conservation assessments, as well as adding older non-digital sources.
ThreatSearch can be used to measure progress toward several targets of the Global Strategy for Plant Conservation. First and foremost, by listing all conservation assessments for plants it helps to track progress towards Target 2 (An assessment of the conservation status of all known plant species, as far as possible, to guide conservation action). ThreatSearch will further help to measure Target 7 (At least 75 per cent of known threatened plant species conserved in situ) and Target 8 (At least 75 per cent of threatened plant species in ex situ collections, preferably in the country of origin, and at least 20 per cent available for recovery and restoration programmes). ThreatSearch also aims to be directly relevant to conservationists, educators, horticulturists, researchers, policy makers and many others who are working to save and understand plant diversity.
GlobalTree Portal
Welcome to the GlobalTree Portal. This portal allows access to information on the world’s nearly 60,000 tree species. On the species pages you can explore tree species distribution, conservation status (global and non-global) and conservation actions. On the country pages you can download a country checklist with associated information on endemism and conservation status. The Global overview allows you to see summary statistics for all trees. The data underlying this portal is information gathered as part of the Global Tree Assessment and links our existing databases GlobalTreeSearch, ThreatSearch, PlantSearch and GardenSearch. In addition, the Conservation Action Tracker enables the monitoring of conservation actions for each tree species. The Conservation Action Tracker can be accessed through individual species pages and you can upload new information through this link.
IWC Population (Abundance) Estimates
Estimates of current population size, known as abundance estimates, are derived from a combination of fieldwork and computer modelling. Vessel-based and aerial sighting surveys, acoustic monitoring, and analysis of individual animal markings are techniques used independently or in conjunction with each other to count whales. The information gathered from this fieldwork is used as the basis for population modelling which produces an abundance estimate.
In 2017, the IWC Scientific Committee established a new expert group to review and agree all the Abundance Estimates submitted to the Scientific Committee. This new process ensures quality and consistency across all the estimates used by the IWC.
Click here to read more about the establishment and work of the Scientific Committee’s Working Group on Abundance Estimates (ASI).
It is impossible for cetacean abundance estimates to be completely precise and IWC estimates are presented as a ‘best estimate’ figure, accompanied by a 95% Confidence Interval, showing a range of plausible values for the population’s actual abundance. As an example, the table below shows that the 1991/92-2003/4 abundance estimate for Southern Hemisphere blue whales is 2,300 with a 95% confidence interval of 1,150-4,500. This means that the range 1,150-4,500 was computed using a method that has a 95% chance of including the population’s actual abundance.
Increases or decreases in population are indicated within the table, where these have been identified.
At present the IWC has agreed estimates only for some species/areas. With the establishment of the ASI group it is planned to update this table and fill in the gaps where possible.
Solutions and case studies
Can we save critically endangered relict endemic plant species? A case study of endemics in Egypt
Rosa arabica and Primula boveana is a perennial endemic to the high mountain area of St. Catherine Protected Area (SCPA) in Egypt and listed as one of the most 100 threatened plants in the world. Recently, they listed as Critically Endangered due to their small Extent of Occurrence and tiny population size (less than 90). The continuous decline in habitat quality for this species and the urgent need to carry out on-ground conservation actions were reported. Many attempts were made in the past to cultivate them in the wild, but they did not succeed. So, this solution aims to conserve them through in situ practices by implementing the following steps, respectively: a) IUCN Red List, b) Ecological Niche Modeling, and c) based on the previous two steps, translocation process for R. arabica in the suitable habitat will be done after carrying out simple layering process (local community traditional method) as one of the most effective traditional vegetative methods for wild cultivation for this species.
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4.1.2. Develop National Red Lists. - es
Primary tools and resources
IUCN Red List index : guidance for national and regional use. Version 1.1
The RLI monitors trends in extinction risk over time and is based on the IUCN Red List of Threatened Species. It tracks the conservation status of major species groups, and has been applied so far to birds, mammals, cycads, amphibians, corals. The data can be disaggregated by region, or country.
How to use
For details of the RLI process and to see results: https://www.iucnredlist.org/assessment/red-list-index
Other tools and resources
National Red List Working Group
The National Red List Working Group (NRLWG) is a working group within the IUCN Red List Committee towards developing and implementing ways to improve linkages between national red listing efforts and the IUCN Red List, and on tools to help national red listing efforts, which are often based on adapting the tools available for global red listing processes to better suit national needs.
The NRLWG also serves as a Coordinating Body for the “National Red List Alliance” (NRLA), established in 2013 to promote and drive forward the national red listing process globally. This was in response to result 2 in the IUCN Red List of Threatened Species Strategic Plan (2013-2020): more IUCN Red List assessments are prepared at national and, where appropriate, at regional scales. The aim is to help countries monitor their progress towards achieving United Nations Sustainable Development Goals (particularly goals 14 and 15) and the Aichi Biodiversity Targets (particularly target 12).
Guidelines for application of IUCN Red List criteria at regional and national levels : version 4.0
The IUCN Red List Categories and Criteria were developed for classifying species at high risk of global extinction, i.e. for assessment at the global level. Guidelines on the application of the IUCN Red List Criteria at national or regional levels were also developed and this volume presents the revised guidelines, published in 2012.
Guidelines for application of IUCN Red list criteria at regional levels, version 3.0
These guidelines provide a logical framework for assessing species at sub-global levels; a means of sharing status information with neighbouring countries; and a suggested format for documentation and publication of listings to improve compatibility with the global IUCN Red List of Threatened Species.
Solutions and case studies
Can we save critically endangered relict endemic plant species? A case study of endemics in Egypt
Rosa arabica and Primula boveana is a perennial endemic to the high mountain area of St. Catherine Protected Area (SCPA) in Egypt and listed as one of the most 100 threatened plants in the world. Recently, they listed as Critically Endangered due to their small Extent of Occurrence and tiny population size (less than 90). The continuous decline in habitat quality for this species and the urgent need to carry out on-ground conservation actions were reported. Many attempts were made in the past to cultivate them in the wild, but they did not succeed. So, this solution aims to conserve them through in situ practices by implementing the following steps, respectively: a) IUCN Red List, b) Ecological Niche Modeling, and c) based on the previous two steps, translocation process for R. arabica in the suitable habitat will be done after carrying out simple layering process (local community traditional method) as one of the most effective traditional vegetative methods for wild cultivation for this species.
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4.1.3. Identify species that require targeted action to enable their recovery. - es
Primary tools and resources
IUCN Red List of Threatened Species
The IUCN Red List of Threatened Species (the Red List) is the globally recognised standard on assessing extinction risk. Red List assessments are used by the Convention on Biological Diversity, CITES, other inter-governmental agreements, national governments, and conservation planners. The Red List assigns species into one of 8 categories based on 5 criteria, all with quantitative thresholds. The three highest categories of threat – Critically Endangered, Endangered, Vulnerable – are collectively considered ‘Threatened’. The Red List process has been formalized to support objectivity and scientific rigour. Red List assessments are carried out by SSC Red List Authorities, Red List Partners, IUCN staff, and regional and national agencies. In addition, Red List assessments provide a compendium of information on species, (taxonomy, geographic range, population, habitat and ecology, threats, conservation actions). The Red List operates at global, regional, and national levels. Guidelines for application at national or regional levels have been developed and the National Red List Working Group provides additional support.
Red List Authorities have been established for all major taxonomic groups included on the IUCN Red List. In most cases, the RLA is a sub-group within an IUCN SSC Specialist Group responsible for the species, groups of species or species within a specific geographic area. There are some exceptions; for example, BirdLife International is the designated RLA for all birds.
The process is guided by Rules of Procedure, supported by guidelines on application of the Red List Categories and Criteria, the Species Information Service (SIS) data entry and management system, and certification for assessors and RL trainers. All these materials can be downloaded, free, from the Red List website. A free, online training course for Red List assessors, is available.
Species can be assessed for the Red List at global, regional, and national levels. Guidelines for the Regional (and national) Application of the Categories and Criteria have been developed and support for development of national Red Lists is available from the National Red List Working Group.
How to use
- To check the global Red List status of a species and access the supporting information, visit the species assessment page on the Red List website
- To download the Red List Categories and Criteria, Guidelines, Rules of Procedure, and Guidelines for Application at Regional Level go the Red List resources
- To initiate or to contribute to a global assessment, contact the Chair of the relevant IUCN SSC Specialist Group or Red List Authority, for birds, see BirdLife International
- For taxonomic groups not listed, contact the relevant IUCN SSC Conservation Committee: Marine, Invertebrate, Plant, Fungi, Freshwater, Red List, Standards and Petitions.
- To develop a National Red List for a species or group of species visit the National Red List Working Group
- To obtain details or register for an online Red List training course: https://www.iucnredlist.org/resources/online
IUCN Green Status of Species
The decline of many species towards extinction has largely focused conservation efforts on ensuring that species remain extant. However, conservationists have long recognised the need to complement this by aiming to recover depleted populations throughout a species’ range and to restore species to ecosystems from which they have been extirpated. The main objectives of the IUCN Green Status of Species are: to provide a standardised framework for measuring species recovery; to recognise conservation achievements; to highlight species whose current conservation status is dependent on continued conservation actions; to forecast the expected conservation impact of planned conservation action; and to elevate levels of ambition for long-term species recovery. These objectives together encourage conservation towards species recovery, throughout a species’ range.
IUCN Red List index : guidance for national and regional use. Version 1.1
The RLI monitors trends in extinction risk over time and is based on the IUCN Red List of Threatened Species. It tracks the conservation status of major species groups, and has been applied so far to birds, mammals, cycads, amphibians, corals. The data can be disaggregated by region, or country.
How to use
For details of the RLI process and to see results: https://www.iucnredlist.org/assessment/red-list-index
Other tools and resources
IWC Extinctions Initiative
According to the IUCN Red List a species or subspecies is considered Extinct (EX) when there is no reasonable doubt that the last individual has died. A species or subspecies is presumed Extinct when exhaustive surveys in known and expected habitat have failed to record a single individual.
Distinct populations can also become locally extinct. A single species is often made up of several separate populations and one population may be eradicated whilst others continue to live. For example, the North Pacific gray whale population in the eastern North Pacific is considered healthy, but the western population is critically endangered.
Extinction is a natural phenomenon, but current rates of extinction are much higher than historic ‘background rates.’ The lifespan of a species varies depending on a number of factors such as body size and geographic range. Many cetacean species are assessed to have survived between 5 and 30 million years. Today’s increased rates of extinction are due to human activities and associated habitat disruption and climate change.
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, assesses the conservation status of most plant and animal species, some subspecies, and some subpopulations. They use a rigorous approach to assess all available data for each species against defined criteria, and classify each into one of the IUCN Red List status categories
IWC Population (Abundance) Estimates
Estimates of current population size, known as abundance estimates, are derived from a combination of fieldwork and computer modelling. Vessel-based and aerial sighting surveys, acoustic monitoring, and analysis of individual animal markings are techniques used independently or in conjunction with each other to count whales. The information gathered from this fieldwork is used as the basis for population modelling which produces an abundance estimate.
In 2017, the IWC Scientific Committee established a new expert group to review and agree all the Abundance Estimates submitted to the Scientific Committee. This new process ensures quality and consistency across all the estimates used by the IWC.
Click here to read more about the establishment and work of the Scientific Committee’s Working Group on Abundance Estimates (ASI).
It is impossible for cetacean abundance estimates to be completely precise and IWC estimates are presented as a ‘best estimate’ figure, accompanied by a 95% Confidence Interval, showing a range of plausible values for the population’s actual abundance. As an example, the table below shows that the 1991/92-2003/4 abundance estimate for Southern Hemisphere blue whales is 2,300 with a 95% confidence interval of 1,150-4,500. This means that the range 1,150-4,500 was computed using a method that has a 95% chance of including the population’s actual abundance.
Increases or decreases in population are indicated within the table, where these have been identified.
At present the IWC has agreed estimates only for some species/areas. With the establishment of the ASI group it is planned to update this table and fill in the gaps where possible.
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4.1.4. Develop comprehensive monitoring programmes, including relevant indicators, for target species and recovery programmes. - es
Primary tools and resources
Biodiversity Indicators Partnership
The Biodiversity Indicators Partnership (BIP) is a global initiative to promote the development, delivery, and use of biodiversity indicators. Its primary role is to delivering indicators to monitor progress toward global and national goals and targets, such as those under the Convention on Biological Diversity, and the Convention on Migratory Species, Ramsar, and others. The BIP also supports indicators used in IPBES Assessment reports and for reporting progress towards Sustainable Development Goals. Indicators supported by the BIP are also used by national and regional governments.
How to use
- To obtain the list of BIP indicators and further guidance: https://www.bipindicators.net/
- To access resources: https://www.bipindicators.net/resources?filters%5Bcategory%5D=47
- To access data by country from the BIP Dashboard, enter the country name or browse by region at: https://bipdashboard.natureserve.org/SelectCountry.html
Database of Global Data Sources for Biodiversity Conservation Monitoring
The database has been developed by the IUCN SSC Species Monitoring Specialist Group with support from Re:wild. It includes 178 global data sources on: monitoring biodiversity state, pressures and threats; monitoring conservation responses to biodiversity loss, and those with multiple uses for biodiversity monitoring. A fact sheet answering frequently asked questions about the database is also available. The Species Monitoring Specialist Group is a voluntary network of experts from around the world and includes specialists on different species and taxa, on different monitoring techniques and issues, and on different types of data use. The group aims to enhance conservation by improving the flow of species data from collection point to decision-maker.
How to use
Version 3.0 of the database and the fact sheet can be downloaded at: https://www.speciesmonitoring.org/data-sources.html
A selection of Manuals, Guidelines and Methods Reviews is available at:
https://www.speciesmonitoring.org/guidelines-and-tools.html
To contact the Species monitoring Specialist Group: SpeciesMonitoring@gmail.com
GEOBON Essential Biodiversity Variables (EBV)
The Group on Earth Biodiversity Observation Network (GEO BON) developed the concept of Essential Biodiversity Variables (EBVs) to advance the collection, sharing, and use of biodiversity information to aggregate, harmonise and interpret biodiversity observations collected by different methods such as in situ monitoring or remote sensing. EBVs can be visualised as biodiversity observations at one location over time, or in many locations, aggregated in a time series of maps. Essential variables to understand climate, biodiversity, and other environmental changes have already been developed (e.g. Essential Climate Variables, Essential Ocean Variables).
How to use
Details of the method, the EBVS already developed, and results are available at: https://geobon.org/ebvs/what-are-ebvs/
Other tools and resources
IUCN SSC Species Monitoring Specialist Group Guidelines and Tools
The IUCN SSC Species Monitoring Specialist Group and the IUCN Global Business and Biodiversity Programme have produced Guidelines for Planning and Monitoring Corporate Biodiversity Performance. The Guidelines were developed by IUCN teams in collaboration with business partners, especially Nespresso, Boskalis and Alcoa.
The Guidelines provide a four-stage, science-based approach for developing a corporate biodiversity strategic plan and measuring biodiversity performance across company operations and supply chains. The approach enables companies to be more targeted in their biodiversity focus by identifying the species and habitats important to them and understanding the benefits they provide to people.
CMP Conservation Standards
The Conservation Standards (CS) are a widely adopted set of principles and practices that bring together common concepts, approaches, and terminology for conservation project design, management, and monitoring. Developed by the Conservation Measures Partnership and regularly updated in collaboration with the broader community, this open-source, strategic process helps conservation teams achieve lasting impact.
Solutions and case studies
Can we save critically endangered relict endemic plant species? A case study of endemics in Egypt
Rosa arabica and Primula boveana is a perennial endemic to the high mountain area of St. Catherine Protected Area (SCPA) in Egypt and listed as one of the most 100 threatened plants in the world. Recently, they listed as Critically Endangered due to their small Extent of Occurrence and tiny population size (less than 90). The continuous decline in habitat quality for this species and the urgent need to carry out on-ground conservation actions were reported. Many attempts were made in the past to cultivate them in the wild, but they did not succeed. So, this solution aims to conserve them through in situ practices by implementing the following steps, respectively: a) IUCN Red List, b) Ecological Niche Modeling, and c) based on the previous two steps, translocation process for R. arabica in the suitable habitat will be done after carrying out simple layering process (local community traditional method) as one of the most effective traditional vegetative methods for wild cultivation for this species.